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A position-dependent silencer plays a major role in repressing. alpha. -fetoprotein expression in human hepatoma

Journal Article · · Molecular and Cellular Biology; (United States)
A large percentage of human hepatomas produce {alpha}-fetoprotein (AFP), but the levels of AFP expression vary greatly among hepatomas. To understand the molecular basis for this variation. The authors analyzed transcriptional regulatory activities associated with the 5{prime}-flanking region of the AFP gene in two human hepatoma cell lines, HuH-7 and huH-1/cl-2, which produce a high and a low level of AFP, respectively. They found that the low level of AFP production in huH-1/cl-2 is due to the action of at least two silencer regions located between the enhancer and the promoter of the AFP gene. In contrast, no silencer activity is expressed in HuH-7. They identified 5{prime}-CTTCATAACTAATACTT-3{prime} to be a core sequence responsible for the negative regulatory activity. This sequence is repeated four times in a strong, distal silencer region, Sd, whereas one copy is present in a weak, proximal silencer region, Sp. The silencer reduces transcriptional initiation by blocking enhancer activation of the AFP promoter in a position-dependent manner. The silencer functions in the presence of positive transcription factors and may play a key role in developmental repression as well as variable expression of the AFP gene in hepatomas.
OSTI ID:
5144644
Journal Information:
Molecular and Cellular Biology; (United States), Journal Name: Molecular and Cellular Biology; (United States) Vol. 11:12; ISSN MCEBD; ISSN 0270-7306
Country of Publication:
United States
Language:
English

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