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Structure of a NEMO/IKK-Associating Domain Reveals Architecture of the Interaction Site

Journal Article · · Structure

The phosphorylation of I{kappa}B by the IKK complex targets it for degradation and releases NF-{kappa}B for translocation into the nucleus to initiate the inflammatory response, cell proliferation, or cell differentiation. The IKK complex is composed of the catalytic IKK{alpha}/{beta} kinases and a regulatory protein, NF-{kappa}B essential modulator (NEMO; IKK{gamma}). NEMO associates with the unphosphorylated IKK kinase C termini and activates the IKK complex's catalytic activity. However, detailed structural information about the NEMO/IKK interaction is lacking. In this study, we have identified the minimal requirements for NEMO and IKK kinase association using a variety of biophysical techniques and have solved two crystal structures of the minimal NEMO/IKK kinase associating domains. We demonstrate that the NEMO core domain is a dimer that binds two IKK fragments and identify energetic hot spots that can be exploited to inhibit IKK complex formation with a therapeutic agent.

Research Organization:
Brookhaven National Laboratory (BNL) National Synchrotron Light Source
Sponsoring Organization:
Doe - Office Of Science
DOE Contract Number:
AC02-98CH10886
OSTI ID:
959992
Report Number(s):
BNL--82978-2009-JA
Journal Information:
Structure, Journal Name: Structure Journal Issue: 5 Vol. 16
Country of Publication:
United States
Language:
English

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