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Competitive Al sup 3+ inhibition of net Mg sup 2+ uptake by intact Lolium multiflorum roots

Journal Article · · Plant Physiology; (USA)
DOI:https://doi.org/10.1104/pp.91.4.1407· OSTI ID:5950965
;  [1]
  1. Louisiana State Univ. Agricultural Center, Baton Rouge (USA)

Aluminum impairs uptake of Mg{sup 2+}, but the mechanisms of this inhibition are not understood. The depletion technique was used to monitor net Mg{sup 2+} uptake from nutrient solution by intact, 23-day-old plants of ryegrass (Lolium multiflorum Lam., cv Gulf and Wilo). Activities of Mg{sup 2+} and monomeric Al species in nutrient solution were calculated and used as the basis for expressing the results. The kinetics of net Mg{sup 2+} absorption was resolved into (a) a transpiration-dependent uptake component, (b) a metabolically mediated, discontinuous saturable component that is Al{sup 3+} sensitive and p-chloromercuribenzene sulfonic acid (PCMBS) resistant, and (c) a linear, carbonyl cyanide m-chlorophenylhydrazone resistant, Al{sup 3+} sensitive component that might be a type of facilitated diffusion. Lowering the pH from 6.0 to 4.2 exerted a noncompetitive inhibition of net Mg{sup 2+} uptake, while aluminum at 6.6 micromolar Al{sup 3+} activity exerted competitive inhibition of net Mg{sup 2+} uptake at pH 4.2. The Al{sup 3+}-induced effect was obvious after 30 minutes. Cultivar-specific ability to retain a higher affinity for Mg{sup 2+} by postulated transport proteins in the presence of Al{sup 3+} might be one of the mechanisms of differential Al tolerance among ryegrass cultivars.

OSTI ID:
5950965
Journal Information:
Plant Physiology; (USA), Journal Name: Plant Physiology; (USA) Vol. 91:4; ISSN 0032-0889; ISSN PLPHA
Country of Publication:
United States
Language:
English