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Fungal community structure and function shifts with atmospheric nitrogen deposition

Journal Article · · Global Change Biology
DOI:https://doi.org/10.1111/gcb.15444· OSTI ID:1808400
 [1];  [2];  [3];  [4];  [4];  [5];  [6];  [7]
  1. Univ. of New Hampshire, Durham, NH (United States); Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States)
  2. Univ. of New Hampshire, Durham, NH (United States); Eidgenoessische Technische Hochschule (ETH), Zurich (Switzerland)
  3. Univ. of New Hampshire, Durham, NH (United States); Univ. of Nebraska, Kearney, NE (United States)
  4. Adam Mickiewicz Univ., Poznan (Poland)
  5. Univ. of New Hampshire, Durham, NH (United States); Young Harris College, GA (United States)
  6. Univ. of Wyoming, Laramie, WY (United States)
  7. Univ. of New Hampshire, Durham, NH (United States)
Fungal decomposition of soil organic matter depends on soil nitrogen (N) availability. This ecosystem process is being jeopardized by changes in N inputs that have resulted from a tripling of atmospheric N deposition in the last century. Soil fungi are impacted by atmospheric N deposition due to higher N availability, as soils are acidified, or as micronutrients become increasingly limiting. Fungal communities that persist with chronic N deposition may be enriched with traits that enable them to tolerate environmental stress, which may trade-off with traits enabling organic matter decomposition. We hypothesized that fungal communities would respond to N deposition by shifting community composition and functional gene abundances toward those that tolerate stress but are weak decomposers. We sampled soils at seven eastern US hardwood forests where ambient N deposition varied from 3.2 to 12.6 kg N ha-1 year-1, five of which also have experimental plots where atmospheric N deposition was simulated through fertilizer application treatments (25–50 kg N ha-1 year-1). Fungal community and functional responses to fertilizer varied across the ambient N deposition gradient. Fungal biomass and richness increased with simulated N deposition at sites with low ambient deposition and decreased at sites with high ambient deposition. Fungal functional genes involved in hydrolysis of organic matter increased with ambient N deposition while genes involved in oxidation of organic matter decreased. One of four genes involved in generalized abiotic stress tolerance increased with ambient N deposition. In summary, we found that the divergent response to simulated N deposition depended on ambient N deposition levels. Fungal biomass, richness, and oxidative enzyme potential were reduced by N deposition where ambient N deposition was high suggesting fungal communities were pushed beyond an environmental stress threshold. Fungal community structure and function responses to N enrichment depended on ambient N deposition at a regional scale.
Research Organization:
Oak Ridge National Laboratory (ORNL), Oak Ridge, TN (United States)
Sponsoring Organization:
National Science Center of Poland; National Science Foundation (NSF); USDOE
Grant/Contract Number:
AC05-00OR22725
OSTI ID:
1808400
Alternate ID(s):
OSTI ID: 1804925
Journal Information:
Global Change Biology, Journal Name: Global Change Biology Journal Issue: 7 Vol. 27; ISSN 1354-1013
Publisher:
WileyCopyright Statement
Country of Publication:
United States
Language:
English

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