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Summary: Cytosystematics of hyperoliid frogs: Phylogeny of Heterixalus, low
karyotypic variability in hyperoliines and separate phylogenetic
position of Leptopelis
G. ODIERNA1
, G. APREA1
, F. ANDREONE2
, W. BOĻ HME3
& M. VENCES4
*
1
Dipartimento di Biologia Funzionale e Strutturale, Universita` di Napoli Federico II, Italy, 2
Museo Regionale di Scienze
Naturali, Torino, Italy, 3
Zoologisches Forschungsinstitut und Museum A. Koenig, Bonn, Germany, and 4
Zoological Institute,
Technical University of Braunschweig, Braunschweig, Germany
Abstract
Karyotypes of 13 species in the anuran family Hyperoliidae are described based on conventional staining, C-banding, Ag-
NOR-banding and staining with fluorochromes (CMA3 and DAPI). The nine studied species of the Malagasy genus
Heterixalus, as well as African species of Acanthixalus, Hyperolius and Kassina, had a karyotype of 2n524 biarmed
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